Throughout the passage of time and the shifts in backgrounds, a particular war continues to leave its mark on history, extending its influence into the modern world. There are two major groups involved in the ensuring war. The first group consists of monsters known as Chiropterans who can change their appearances into that of human beings. They are actually immortals who feed on the blood of the living. The second group is an organisation known as the "Red Shield", formed to track down these monsters and exterminate them. Otonashi Saya is a high-school girl who lives a peaceful life with her family. The only problem is, she has no recollections of her life beyond that of the past year. However, her happy life will be destroyed by an event that will lead her to her destined fate.
Characters
Chiroptera
Chiroptera are hematophagous bat-like creatures that live by feeding on human blood. They generally possess extraordinary speed, strength, and supernatural healing abilities. They are also capable of contorting and transforming various body parts in order to fly or have longer limbs and higher level Chiroptera have the ability to disguise themselves as ordinary humans.
As Chiroptera can heal almost instantly from most wounds, bullets are non-lethal and only useful in slowing them down. Even explosions do not seem to be capable of killing them, though if buried or sealed inside stone they will go into a sleep state until freed.
Chiroptera are created from the blood of a Queen, and ironically it is also the primary method of killing them. When the blood of the other Queen is introduced into a Chiropteran's bloodstream, it almost instantly crystallizes and destroys them. So Chiroptera created from Diva's blood can be killed by Saya's blood, and visa versa.
Through human experimentation with Diva's blood, a substance called "Delta 67" was created. When "Delta 67" is introduced into the blood stream of humans, they change into Chiroptera. As they are all formed from Diva's blood, they can be killed through the induction of Saya's blood.
Artificial chiropterans
The most basic form of Chiropterans that are created from the straight infusion of Delta 67 into a human host. The human host quickly loses their human intelligence and memories, transforming into a form resembling a horse-headed gorilla with fangs. Phase 1 Chiropterans live and act purely on instinct, hunting and killing humans, and sometimes animals, for their blood.
Through the research efforts of Dr. Boris and Amshel, a higher level artificial Chiroptera was created from the Delta 67 which were collectively called the Schiff. Unlike Phase 1 Chiropterans, the Schiff have human level intelligence and emotions, though somewhat stunted by having lived most of their lives caged. They are aggressive, but this is due to their being trained as warriors. As they are exposed to other ways of living after escaping the lab, they seem to be able to learn and adapt.[6] The Schiff are unique among the Chiropterans as they can be killed by exposure to direct sunlight. They also suffer from a condition called Thorn, which slowly crystallizes and kills them.
Using the genetic template of Moses, the leader of the Schiff, an improved version of the Schiff were created called the Corpse Corps. Like the Schiff, the Corpse Corps are capable fighters with bladed weapons and are marketed to various national military organizations as "Chiropteran exterminators", despite being Chiropterans themselves.
Chevaliers
Chevaliers are the second highest level of Chiroptera in terms of strength and ability. They are created when a human drinks the blood of a Queen. This creates a powerful bond between the Chevalier and the Queen that sired them, and they are mentally compelled to serve and protect her at all costs.
Chevaliers have even greater enhanced speed, strength and endurance than regular Chiroptera. Though they retain the appearance in body form and age that they had when they were transformed, Chevaliers have vast shape-shifting abilities. They can transform into unique forms resembling regular Chiroptera, either entirely or limited to specific body parts, and can transform into the humans they have fed upon. They can instantly heal from almost any non-lethal wound, though severe wounds will temporarily cause an increase in their need for blood.
Queens
Queens, also called the originals, are the first Chiroptera. In any given generation, there are always two Queens, who are female twins born from one of the last Queens, and they are naturally antagonistic with one another. Saya and Diva are the current Queens of their species. They were discovered by Joel Goldschmidt in the 19th century inside a mysterious creature found in Iceland known as SAYA. As part of his experiment, he raised Saya like a daughter, but kept Diva locked away in a tower.
All other Chiroptera are born of the blood of Queens, either through the Queen's own efforts or human experimentation. Unlike all other Chiroptera, Queens appear to retain a human appearance at all times, though they can change their human appearances somewhat and their mother's corpse had a Chiropteran appearance. Their blood can also crystallize and destroy any Chiroptera of any level that was created by their sister Queen.
Queens spend only a few years awake at any given time, between decades of hibernation inside a cocoon. Queens have enhanced speed, strength, and endurance that surpasses those of their Chevaliers and other Chiroptera. They may also have some shape-shifting abilities, though during the series their abilities seem to be limited to modifying to their outward human appearance. The strength of a Queen's traits are dependent on their feeding regularly on human blood. As such, Diva, who freely feeds on humans, can also freely use all of her abilities, while Saya, who lives almost entirely on transfusions, is usually only slightly above human level strength.
When a Queen becomes pregnant with the next generation of Queens, her blood will lose its dangerous qualities. A Queen can only be impregnated by a Chevalier created by her sister Queen, insuring the purity of the species is maintained.
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